Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale. It was "cloned" from a model that InsightMaker provides to its users, at
https://insightmaker.com/insight/2068/Isle-Royale-Predator-Prey-Interactions
Thanks Scott Fortmann-Roe.

I've created a Mathematica file that replicates the model, at
http://www.nku.edu/~longa/classes/2018spring/mat375/mathematica/Moose-n-Wolf-InsightMaker.nb

It allows one to experiment with adjusting the initial number of moose and wolves on the island.

I used steepest descent in Mathematica to optimize the parameters, with my objective data being the ratio of wolves to moose. You can try my (admittedly) kludgy code, at
http://www.nku.edu/~longa/classes/2018spring/mat375/mathematica/Moose-n-Wolf-InsightMaker-BestFit.nb

{WolfBirthRateFactorStart,
WolfDeathRateStart,
MooseBirthRateStart,
MooseDeathRateFactorStart,
moStart,
woStart} =
{0.000267409,
0.239821,
0.269755,
0.0113679,
591,
23.};

Clone of Isle Royale: Predator/Prey Model for Moose and Wolves
Insight diagram
Clone of Lab 2 Part 3
Insight diagram

This model has two main components. First is modelling the change in population composition as non-First Nations immigration increases with the opening of new mines in the region. The second is modelling the increasing income disparity between First Nations and non-First Nations as mining jobs are disproportionately gained by non-First Nations workers.

Clone of Northern Ontario Demographic and Income Trend Model
Insight diagram
Simple BIDE model
Clone of Population - BIDE
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
This simulation shows how plant, deer and wolf populations impact each other in a deciduous forest ecosystem.
Clone of Plant, Deer and Wolf Population Dynamics G-IV Intro
Insight diagram

Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")
Tags: Education, Chaos, Ecology, Biology, Population
Thanks to Insight Author: John Petersen

Edits by Andy Long

Everything that follows the dashes was created by John Petersen (or at least came from his Insight model). I just wanted to make a few comments.

We are looking at Hare and Lynx, of course. Clone this insight, and change the names.

Then read the text below, to get acquainted with one of the most important and well-known examples of a simple system of differential equations in all of mathematics.

http://www.nku.edu/~longa/classes/mat375/mathematica/Lotka-Volterra.nb
------------------------------------------------------------

Dynamic simulation modelers are particularly interested in understanding and being able to distinguish between the behavior of stocks and flows that result from internal interactions and those that result from external forces acting on a system. 

For some time modelers have been particularly interested in internal interactions that result in stable oscillations in the absence of any external forces acting on a system. 

The model in this last scenario was independently developed by Alfred Lotka (1924) and Vito Volterra (1926).  Lotka was interested in understanding internal dynamics that might explain oscillations in moth and butterfly populations and the parasitoids that attack them.  Volterra was interested in explaining an increase in coastal populations of predatory fish and a decrease in their prey that was observed during World War I when human fishing pressures on the predator species declined. 

Both discovered that a relatively simple model is capable of producing the cyclical behaviors they observed. 

Since that time, several researchers have been able to reproduce the modeling dynamics in simple experimental systems consisting of only predators and prey.  It is now generally recognized that the model world that Lotka and Volterra produced is too simple to explain the complexity of most predator-prey dynamics in nature.  And yet, the model significantly advanced our understanding of the critical role of feedback in predator-prey interactions and in feeding relationships that result in community dynamics.

The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.

As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of MAT375 Clone of Bio103 Predator-Prey Model ("Lotka'Volterra")
Insight diagram
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

Experiment with adjusting the moose birth-rate to simulate Over-shoot followed by environmental recovery
Clone of Clone of Royal Island- Resilience
Insight diagram
Overview
A model which simulates the competition between logging versus adventure tourism (mountain bike ridding) in Derby Tasmania.  Simulation borrowed from the Easter Island simulation.

How the model works.
Trees grow, we cut them down because of demand for Timber amd sell the logs.
With mountain bkie visits.  This depends on past experience and recommendations.  Past experience and recommendations depends on Scenery number of trees compared to visitor and Adventure number of trees and users.  Park capacity limits the number of users.  
Interesting insights
It seems that high logging does not deter mountain biking.  By reducing park capacity, visitor experience and numbers are improved.  A major problem is that any success with the mountain bike park leads to an explosion in visitor numbers.  Also a high price of timber is needed to balance popularity of the park. It seems also that only a narrow corridor is needed for mountain biking
Clone of Simulation of Derby Mountain biking versus logging
Insight diagram
Start with logistic population dynamics (which can't overshoot) but then add delay in the "feedback signal" (the approach to the carrying capacity). One species, able to exploit one resource, which is available at  a fixed, finite, flow (not a depleting stock). At low populations, growth is exponential. As long as population below carrying capacity, growth continues. Without delay, it will smoothly stabilize at the carrying capacity. But with delay, it will overshoot; but oscillation should dampen, so eventually still stabilizes. Similar dynamics. "from above" (if, e.g., "initial" population somehow above carrying capacity; or, more plausibly, if carrying capacity dynamically falls to some lower level). With more delay, get more extreme overshoot. In "extreme" cases (relatively large delay, large overshoot) we can note asymmetry in "boom" and "bust" - bust is more rapid. This can be interpreted as a very simple version of Bardi's Seneca Cliff.
Population dynamics with overshoot ("Seneca cliff"?)
Insight diagram


Clone of Prey&Predator
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram

Addition of Allee effect to Logistic Growth Insight 1540. From the Appendix of Marten Scheffer's 2009 Book Critical Transitions in Nature and Society p332 http://bit.ly/yrd3GN. The Allee affect describes a threshold density (smaller than the carrying capacity K) below which populations go into free fall extinction.

Allee Effect from Critical Transitions
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Clone of Prey&Predator
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale. It was "cloned" from a model that InsightMaker provides to its users, at
https://insightmaker.com/insight/2068/Isle-Royale-Predator-Prey-Interactions
Thanks Scott Fortmann-Roe.

I've created a Mathematica file that replicates the model, at
http://www.nku.edu/~longa/classes/2018spring/mat375/mathematica/Moose-n-Wolf-InsightMaker.nb

It allows one to experiment with adjusting the initial number of moose and wolves on the island.

I used steepest descent in Mathematica to optimize the parameters, with my objective data being the ratio of wolves to moose. You can try my (admittedly) kludgy code, at
http://www.nku.edu/~longa/classes/2018spring/mat375/mathematica/Moose-n-Wolf-InsightMaker-BestFit.nb

{WolfBirthRateFactorStart,
WolfDeathRateStart,
MooseBirthRateStart,
MooseDeathRateFactorStart,
moStart,
woStart} =
{0.000267409,
0.239821,
0.269755,
0.0113679,
591,
23.};

Clone of Isle Royale: Predator/Prey Model for Moose and Wolves
Insight diagram
Huemul-ciervo_colorado(sin_caza)-Ks_constantes
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator
Insight diagram
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

We incorporate logistic growth into the moose dynamics, and we replace the death flow of the moose with a kill rate modeled from the kill rate data found on the Isle Royale website.

I start with these parameters:
Wolf Death Rate = 0.15
Wolf Birth Rate = 0.0187963
Moose Birth Rate = 0.4
Carrying Capacity = 2000
Initial Moose: 563
Initial Wolves: 20

I used RK-4 with step-size 0.1, from 1959 for 60 years.

The moose birth flow is logistic, MBR*M*(1-M/K)
Moose death flow is Kill Rate (in Moose/Year)
Wolf birth flow is WBR*Kill Rate (in Wolves/Year)
Wolf death flow is WDR*W

Clone of Final Midterm Student version of A More Realistic Model of Isle Royale: Predator Prey Interactions
Insight diagram
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


Clone of Prey&Predator