​Physical meaning of the equations  The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:        1. The prey population finds ample food at all times.    2. The food supply of the predator population depends entirely on the
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


 ​Physical meaning of the equations  The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:        1. The prey population finds ample food at all times.    2. The food supply of the predator population depends entirely on the
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.  Experiment with adjusting the initial number of moose and wolves on the island.
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

Experiment with adjusting the initial number of moose and wolves on the island.
 This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.    Reference:  Hartig, F., Minunno, F., and Paul, S. (2019). Baye
This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.

Reference:
Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
 This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.    Reference:  Hartig, F., Minunno, F., and Paul, S. (2019). Baye
This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.

Reference:
Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
 ​Physical meaning of the equations  The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:        1. The prey population finds ample food at all times.    2. The food supply of the predator population depends entirely on the
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


 STEM-SM combines a simple ecosystem model (modified version of VSEM; Hartig et al. 2019) with a soil moisture model (Guswa et al. (2002) leaky bucket model). Outputs from the soil moisture model influence ecosystem dynamics in three ways.   (1) The ratio of actual transpiration to maximum evapotran
STEM-SM combines a simple ecosystem model (modified version of VSEM; Hartig et al. 2019) with a soil moisture model (Guswa et al. (2002) leaky bucket model). Outputs from the soil moisture model influence ecosystem dynamics in three ways. 
(1) The ratio of actual transpiration to maximum evapotranspiration (T/ETmax) modifies gross primary productivity (GPP).
(2) Degree of saturation of the soil (Sd) modifies the rate of soil heterotrophic respiration.
(3) Water limitation of GPP (by T/ETmax) and of soil nutrient availability (approximated by Sd) combine with leaf area limitation (approximated by fraction of incident photosynthetically-active radiation that is absorbed) to modify the allocation of net primary productivity to aboveground and belowground parts of the vegetation.

Ecosystem dynamics in turn influence flows of water in to and out of the soil moisture stock. The size of the aboveground biomass stock determines fractional vegetation cover, which modifies interception, soil evaporation and transpiration by plants.

References:
Guswa, A.J., Celia, M.A., Rodriguez-Iturbe, I. (2002) Models of soil moisture dynamics in ecohydrology: a comparative study. Water Resources Research 38, 5-1 - 5-15.

Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools

11 months ago
 This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.    Reference:  Hartig, F., Minunno, F., and Paul, S. (2019). Baye
This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.

Reference:
Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
​Modelo retirado do link  https://insightmaker.com/insight/71649/Fern-Population-Model
​Modelo retirado do link 
https://insightmaker.com/insight/71649/Fern-Population-Model
 STEM-SM combines a simple ecosystem model (modified version of VSEM; Hartig et al. 2019) with a soil moisture model (Guswa et al. (2002) leaky bucket model). Outputs from the soil moisture model influence ecosystem dynamics in three ways.   (1) The ratio of actual transpiration to maximum evapotran
STEM-SM combines a simple ecosystem model (modified version of VSEM; Hartig et al. 2019) with a soil moisture model (Guswa et al. (2002) leaky bucket model). Outputs from the soil moisture model influence ecosystem dynamics in three ways. 
(1) The ratio of actual transpiration to maximum evapotranspiration (T/ETmax) modifies gross primary productivity (GPP).
(2) Degree of saturation of the soil (Sd) modifies the rate of soil heterotrophic respiration.
(3) Water limitation of GPP (by T/ETmax) and of soil nutrient availability (approximated by Sd) combine with leaf area limitation (approximated by fraction of incident photosynthetically-active radiation that is absorbed) to modify the allocation of net primary productivity to aboveground and belowground parts of the vegetation.

Ecosystem dynamics in turn influence flows of water in to and out of the soil moisture stock. The size of the aboveground biomass stock determines fractional vegetation cover, which modifies interception, soil evaporation and transpiration by plants.

References:
Guswa, A.J., Celia, M.A., Rodriguez-Iturbe, I. (2002) Models of soil moisture dynamics in ecohydrology: a comparative study. Water Resources Research 38, 5-1 - 5-15.

Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools

Challenges in sustainability are multilevel. This diagram attempts to summarize levels of self reinforcing destructive dynamics, authors that deal with them, and point of leverage for change.  The base of the crisis is a mechanistic rather than ecological worldview. This mechanistic worldview is bas
Challenges in sustainability are multilevel.
This diagram attempts to summarize levels of self reinforcing destructive dynamics, authors that deal with them, and point of leverage for change.

The base of the crisis is a mechanistic rather than ecological worldview. This mechanistic worldview is based on outdated science that assumed the universe to be a large machine. In a machine there is an inside and an outside. The health of the inside is important for the machine, the outside not. In an ecological view everything is interconnected, there is no clear separation in the future of self and other. All parts influence the health of other parts. To retain health sensitivity and democracy are inherent. The sense of separation from other that keeps the mechanistic worldview dominant is duality. Being cut off from spiritual traditions due to a mechanistic view of science people need access to inter-spirituality to reconnect with the human traditions and tools around connectedness, inner discovery, and compassion. Many books on modern physics and biology deal with the system view implications. "The coming interspiritual age" deals with the need to connect spiritual traditions and science.

At the bottom for the dynamic is an individual a sense of disconnectedness leads to a dependency on spending and having rather than connecting. The connecting has become too painful and dealing with it unpopular in our culture. Joanna Macy deals with this in Active Hope. 

This affluenza and disconnection is worsened by a market that floods one with advertisements aimed at creating needs and a sense of dissatisfaction with that one has.

National economies are structured around maximising GDP which means maximising consumption and financial capital movement. This is at the cost of local economies. These same local economies are needed for balanced happiness as well as for sustainability.

Generally institutions focus on maximising consumption rather than sustaining life support systems. David Korten covers this well.

Power and wealth is confused in this worldview. In striving for wealth only power is striven for in the form of money and monopoly.

Those at the head of large banks and corporations tend to be there because they exemplify this approach. They have few scruples about enforcing this approach onto everyone through wars and disaster capitalism. Naomi Klein and David Estulin documented this.

Power has become so centralized that we need this understanding to be widespread and include many of those in power. Progress of all of these levels are needed to show them and all that another way is possible.
 ​Physical meaning of the equations  The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:        1. The prey population finds ample food at all times.    2. The food supply of the predator population depends entirely on the
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale. It was "cloned" from a model that InsightMaker provides to its users, at  https://insightmaker.com/insight/2068/Isle-Royale-Predator-Prey-Interactions  Thanks Scott Fortmann-Roe.
This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale. It was "cloned" from a model that InsightMaker provides to its users, at
https://insightmaker.com/insight/2068/Isle-Royale-Predator-Prey-Interactions
Thanks Scott Fortmann-Roe.

I've created a Mathematica file that replicates the model, at
http://www.nku.edu/~longa/classes/2018spring/mat375/mathematica/Moose-n-Wolf-InsightMaker.nb

It allows one to experiment with adjusting the initial number of moose and wolves on the island.

I used steepest descent in Mathematica to optimize the parameters, with my objective data being the ratio of wolves to moose. You can try my (admittedly) kludgy code, at
http://www.nku.edu/~longa/classes/2018spring/mat375/mathematica/Moose-n-Wolf-InsightMaker-BestFit.nb

{WolfBirthRateFactorStart,
WolfDeathRateStart,
MooseBirthRateStart,
MooseDeathRateFactorStart,
moStart,
woStart} =
{0.000267409,
0.239821,
0.269755,
0.0113679,
591,
23.};

 ​Physical meaning of the equations  The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:        1. The prey population finds ample food at all times.    2. The food supply of the predator population depends entirely on the
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


 ​Physical meaning of the equations  The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:        1. The prey population finds ample food at all times.    2. The food supply of the predator population depends entirely on the
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


 ​Physical meaning of the equations  The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:        1. The prey population finds ample food at all times.    2. The food supply of the predator population depends entirely on the
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


 ​Physical meaning of the equations  The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:        1. The prey population finds ample food at all times.    2. The food supply of the predator population depends entirely on the
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


 ​Physical meaning of the equations  The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:        1. The prey population finds ample food at all times.    2. The food supply of the predator population depends entirely on the
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.


 This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.    Reference:  Hartig, F., Minunno, F., and Paul, S. (2019). Baye
This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.

Reference:
Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
 This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.    Reference:  Hartig, F., Minunno, F., and Paul, S. (2019). Baye
This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.

Reference:
Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
 This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.  Experiment with adjusting the initial number of moose and wolves on the island.

This model illustrates predator prey interactions using real-life data of wolf and moose populations on the Isle Royale.

Experiment with adjusting the initial number of moose and wolves on the island.

This is a model which explains the difference between Mountain bikes riding compared to logging in the Tasmanian forests.
This is a model which explains the difference between Mountain bikes riding compared to logging in the Tasmanian forests.
 This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.    Reference:  Hartig, F., Minunno, F., and Paul, S. (2019). Baye
This model is a modified version of the 'Very Simple Ecosystem Model' (VSEM; Hartig et al. 2019). Controls have been added to gross primary productivity (GPP) and heterotrophic respiration (Rhetero) based on evapotranspiration rates.

Reference:
Hartig, F., Minunno, F., and Paul, S. (2019). BayesianTools: General-Purpose MCMC and SMC Samplers and Tools for Bayesian Statistics. R package version 0.1.7. https://CRAN.R-project.org/package=BayesianTools
 ​Physical meaning of the equations  The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:        1. The prey population finds ample food at all times.    2. The food supply of the predator population depends entirely on the
​Physical meaning of the equations
The Lotka–Volterra model makes a number of assumptions about the environment and evolution of the predator and prey populations:

1. The prey population finds ample food at all times.
2. The food supply of the predator population depends entirely on the size of the prey population.
3. The rate of change of population is proportional to its size.
4. During the process, the environment does not change in favour of one species and genetic adaptation is inconsequential.
5. Predators have limitless appetite.
As differential equations are used, the solution is deterministic and continuous. This, in turn, implies that the generations of both the predator and prey are continually overlapping.[23]

Prey
When multiplied out, the prey equation becomes
dx/dtαx - βxy
 The prey are assumed to have an unlimited food supply, and to reproduce exponentially unless subject to predation; this exponential growth is represented in the equation above by the term αx. The rate of predation upon the prey is assumed to be proportional to the rate at which the predators and the prey meet; this is represented above by βxy. If either x or y is zero then there can be no predation.

With these two terms the equation above can be interpreted as: the change in the prey's numbers is given by its own growth minus the rate at which it is preyed upon.

Predators

The predator equation becomes

dy/dt =  - 

In this equation, {\displaystyle \displaystyle \delta xy} represents the growth of the predator population. (Note the similarity to the predation rate; however, a different constant is used as the rate at which the predator population grows is not necessarily equal to the rate at which it consumes the prey). {\displaystyle \displaystyle \gamma y} represents the loss rate of the predators due to either natural death or emigration; it leads to an exponential decay in the absence of prey.

Hence the equation expresses the change in the predator population as growth fueled by the food supply, minus natural death.